The index is most often used for ecological studies that measure species diversity, but the same analysis can also be applied to other principles, such. It measures the rarity and commonness of species in a community. Wiener index, shannonweaver index, species richness, species diversity. The shannon index measures the order within a community written as h. Which of these diversity indices is the basic shannon index. In statistical thermodynamics the most general formula for the thermodynamic entropy s of a thermodynamic system is the gibbs entropy.
An index of species diversity devised in 1949 by e. But other forces greater than himself are at play, and the multilayered, complex, and nuanced on the surface, this novel tells the story of robert shannon, a former chaplain with the u. Of the many species diversity indices used in the literature, the shannon index is perhaps most commonly used. A diversity index is a quantitative measure that reflects the number of different. If we use the compliment to simpson s d, the value is. Thus, if sunflower city was 85% white and 15% black, the index of diversity would be. Species diversity species diversity differs from species richness in that it takes into account both the numbers of species present and the dominance or evenness of species in relation to one another. The higher the value of this inverse index the greater the diversity. The shannon index increases as both the richness and the evenness of the community increase. First, let us calculate the sum of the given values. Like simpsons index, shannon s index accounts for both abundance and evenness of the species present. It is calculated by dividing the shannon diversity index by its maximum h m.
A tribute to claude shannon 19162001 and a plea for. Diversity index analysis tool real statistics using excel. Shannonwiener diversity index is a term in biologyecology. Diversity indices that combine both richness and abundance i. A diversity index is the measure of species diversity in a given community. Shannon index the idea behind this index is that the diversity of a community is similar to the amount of information in a code or message.
B10 of figure 1 of shannons diversity index except that the indices must be multiplied by 2. Here is the website i show you toward the end of the. Divide shannons diversity index h by natural logarithm of species richness lns to calculate the species evenness. An example of the use of the index of diversity would be a measure of racial diversity in a city. The methods on offer range from the familiar, such as the calculation of various diversity indices and the fitting of common distributions, to more recentlydeveloped techniques such as total species complement estimators. Introduction, simpsons index and shannonweiner index. Larger values reflect communities with greater species richness and evenness, while lower numbers reflect communities with fewer species andor a very uneven distribution. Pdf application of shannons index to study diversity with. Shannon s index of diversity h is derived from information theory, originally in the context of information in telephone systems shannon, 1948.
Aug 28, 2014 for example, when modeling changes in species diversity throughout a community, knowledge of the genetic and chemical diversity of the primary producer e. The shannonweiner index is most sensitive to the number of species in a sample, so it is usually considered to be biased toward measuring species richness. Note that species evenness ranges from zero to one, with zero signifying no evenness and one, a complete evenness. How can i calculate shannons diversity index using excel sheet. A diversity index is a quantitative measure that reflects how many different types such as. May 18, 2018 here i show how to use the shannon weiner diversity index in order to describe the variety of organisms. Simpsons diversity index in a dictionary of ecology. If the community was equitable first example, wed have a 5050 chance of sampling the first species. Find shannons index of diversity and index of relative diversity for a random sample of 25 observations distributed among five categories as shown in range b4. Here i show how to use the shannonweiner diversity index in order to describe the variety of organisms. It is different from species richness in that unlike richness it also shows community composition and takes into account the relative abundance of species that are present in the community. Diversity indices like the shannon entropy shannon wiener index and the ginisimpson index are not themselves diversities. Shannon wiener species diversity index formula data analysis. Species diversity is also described in terms of the phylogenetic diversity, or evolutionary relatedness, of the species present in an area.
The inspiration for adopting the word entropy in information theory came from the close resemblance between shannon s formula and very similar known formulae from statistical mechanics. We will then consider one measure used to compare diversity among communities, jaccard. It was the result of crucial contributions made by many distinct individuals, from a variety of backgrounds, who took his ideas and expanded upon them. Information theory was not just a product of the work of claude shannon.
The simpson index is a dominance index because it gives more weight to common or dominant species. This is following the method proposed by jost et al. Natural resource biometrics measures of diversity shannon. In fact, in the late 1940s shannon had built on the work of wiener.
A diversity index also called phylogenetic indices or phylogenetic metrics is a quantitative measure that reflects how many different types such as species there are in a dataset a community and that can simultaneously take into account the phylogenetic relations among the individuals distributed among those types, such as richness, divergence or evenness. This diversity measure is based on information theory. Diversity indices like the shannon entropy shannonwiener index and the ginisimpson index are not themselves diversities. As a measure of species diversity, we will calculate the shannon index. Diversity of classes shannon, simpson, species statsdirect. Shannon is sent to ireland in the summer of 1922, supposedly to trace his familys roots along the shannon river.
In this example, the first sample would be considered more diverse. How do i run diversity index calculation on indivi. Jan 26, 2012 a diversity index is the measure of species diversity in a given community. Sample values s 60,10,25,1,4 number of species n 5. Sample values s 60,10,25,1,4 number of species n 5 first, let us calculate the sum of the given values. For our uses, this order could be characterized by the number of species andor the number of individuals in each species, within our sample plot.
Addition of menhinick, gini and strongs alpha diversity indices. Indeed the diversity and directions of their perspectives and interests shaped the direction of information theory. The shannonweaver diversity index is based on communication theory. The same process operates with the shannon index which should anyway be replaced with exponential shannon one of the hill numbers and other diversity measures, though the problem has too. Code to add this calci to your website just copy and paste the below code to your webpage where you want to display this calculator. Just like the diameter of a sphere is an index of its. Shannon in 1949 derived a function that became known as the shannon diversity index. In the shannon index, p is the proportion nn of individuals of one particular species found n divided by the total number of individuals found n, ln is the natural log. The fact that the index incorporates both components of biodiversity can be seen as both a strength and a weakness. The shannon diversity index is a commonly used measure of diversity. The data are collected from the statistical hand book, assam, 2004 1. A greater number of species and a more even distribution both increase diversity as measured by h.
How to do the shannonweiner diversity calculation youtube. For example, species richness usually noted s of a dataset is the number of different. The document is still incomplete and does not cover all diversity methods in vegan. It is characterized by the number of individuals observed for each species in the ecosystem.
Like simpsons index, shannons index accounts for both abundance and evenness of the species present. Arguments data a list of otu tables to be processed. Shannons index of diversity h is derived from information theory, originally in the context of information in telephone systems shannon, 1948. What is the correct name for the biodiversity index. Shannon index is a commonly used diversity index that takes into account.
In this second example, if we were to sample 1 individual, it likely would be of the first species. A tribute to claude shannon 19162001 and a plea for more rigorous use of species richness, species diversity and the shannon wiener index. The resulting species abundance distribution, the log series model, is associated with an index of diversity known as fishers other models and measures were soon developed, and two enduring approaches are prestons log normal distribution of species abundances and. The concept of information entropy was introduced by claude shannon in his 1948 paper a mathematical theory of communication. It is a transmission model consisting of five elements.
Richness r simply quantifies how many different types the dataset of interest contains. Oxford university press usa publishes scholarly works in all academic disciplines, bibles, music, childrens books, business books, dictionaries, reference books, journals, text books and more. Three commonly used alpha diversity measures include species observed, shannon index and simpson index. Feb 12, 2016 shannonwiener diversity index in excel duration. The calculator uses the following formula to calculate the shannonwiener diversity index. Shannon wiener species diversity index calculator calculate. The one formula that i have considered is the shannon index, but that measures the diversity with more than one species in a community, not the species attribute to the community. It covers the number of different land cover types m observed along the straight line and their relative abundances p i.
H has no intuitive interpretation in terms of probability and is sensitive to sample size. For example, species richness usually noted s of a dataset is the number of different species in the corresponding species list. As has been noted previously by some authors for example. Richness is a simple measure, so it has been a popular diversity index in ecology, where abundance data are often not available for the datasets of interest. Calculate the shannon diversity index and evenness for these sample values. On some occasions it is called the shannonwiener index and on other occasions it is called the shannonweaver index. The hutcheson ttest is a modified version of the classic ttest that provides a way to compare two samples. Application of shannons index to study diversity with. Simpson, given by d1pi2, where d is the diversity index and p i is the proportion of individuals in the ith species.
Statistics shannon wiener diversity index tutorialspoint. Foremost among these are the shannons diversity h and simpsons diversity d 1 indices table table1, 1, which differ in their theoretical foundation and interpretation magurran 2004. Shannons diversity index real statistics using excel. Shannon by frank delaney meet your next favorite book. The proportion of species i relative to the total number of species p i is calculated, and then multiplied by the natural logarithm of this proportion lnp i. The shannonwiener diversity index, h, is calculated using the following equation. Popular diversity books meet your next favorite book. The most wellknown and influential formal model of communication, developed in 1949 by claude shannon and warren weaver see communication models.
How to calculate shannon wiener diversity index definition. It is not itself diversity, but just indices of diversity. Calculations of shannon diversity index suggest that individuals enter the sample randomly from an. What does the diversity index take into account when measuring diversity. The uncertainty is measured by the shannon function h. The first step in calculating the shannon diversity index of your closet is counting how many species life forms you have. The shannon diversity index h is another index that is commonly used to characterize species diversity in a community. Shannonweiner index the shannon index is affected by both the number of species and their equitability, or evenness. Species diversity considers both the number of species richness in the community and their relative abundance evenness or equability. Since by definition the pis will all be between zero and one, the. Lets compute the shannonweiner diversity index for the same hypothetical community in the previous example. H has its foundations in information theory and represents the uncertainty about the. There have been numerous attempts to create compound indices that combine measures of richness and abundance.
Species diversity and richness 4 is designed for professional ecologists and their students. The information entropy, often just entropy, is a basic quantity in information theory associated to any random variable, which can be interpreted as the average level of information, surprise, or uncertainty inherent in the variables possible outcomes. The maximum diversity hmax of a sample is found when all species are equally abundant. Online biodiversity calculator which helps to calculates the shannon s diversity index and evenness from the given sample data. Simpson, given by d1pi2, where d is the diversity index and pi is the proportion of individuals in the ith species. For example, some areas may be rich in closely related taxa, having evolved from a common ancestor that was also found in that same area, whereas other areas may have an array of less closely related. However, you cannot compare the two index values using classic hypothesis tests because you do not have replicated data. It combines both evenness and richness in a single measure. When calculating the shannon index is often used a binary logarithm, but also.
A diversity index is a mathematical measure of species diversity in a community. Shannon weiner index the shannon index is affected by both the number of species and their equitability, or evenness. It is often used when we wish to study a random sample from a larger community. Here p i is the proportion of total number of species made up of the ith. Diversity indices provide more information about community composition than simply species richness i. They are just indices of diversity, in the same way that the diameter of a sphere is an index of its volume but is not itself the volume.
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